When Ugly Betty first started, we knew editor of Mode magazine - Daniel Meade's - face looked familiar...

Then it clicked... he played the evil Dean in the O.C. who had an affair with Taylor! Check him out at the end of the video below:

Not only that, he played closet homosexual Greg in the retro teen flick Cruel Intentions - sharing a bed with none other than gorgeous Joshua Jackson!! Here they are in a still shot from the film:

Eric and Josh in Cruel Intentions

This is how you'll recognize him now though, as Daniel Meade in Ugly Betty:

Eric and America Ferrera in a scene from Ugly Betty

Did you recognize him too?!